diff --git a/pkl-core/src/main/java/org/pkl/core/ast/builder/AstBuilder.java b/pkl-core/src/main/java/org/pkl/core/ast/builder/AstBuilder.java
index d7df51d14..dee0f7926 100644
--- a/pkl-core/src/main/java/org/pkl/core/ast/builder/AstBuilder.java
+++ b/pkl-core/src/main/java/org/pkl/core/ast/builder/AstBuilder.java
@@ -724,7 +724,7 @@ public class AstBuilder extends AbstractAstBuilder<Object> {
       //     elem
       //   }
       // }
-      return p.levelsUp() == 0
+      return p.levelsUp() == 0 && !p.needsFrameSkip()
           ? ReadExactFrameSlotNodeGen.create(sourceSection, p.slot())
           : ReadFrameSlotNodeGen.create(
               sourceSection, p.slot(), new GetEnclosingFrameNode(p.levelsUp()));
diff --git a/pkl-core/src/main/java/org/pkl/core/ast/builder/SymbolTable.java b/pkl-core/src/main/java/org/pkl/core/ast/builder/SymbolTable.java
index 762af9de0..6dccb304b 100644
--- a/pkl-core/src/main/java/org/pkl/core/ast/builder/SymbolTable.java
+++ b/pkl-core/src/main/java/org/pkl/core/ast/builder/SymbolTable.java
@@ -509,6 +509,7 @@ public final class SymbolTable {
     private @Nullable <R> R resolveLexical(ResolutionFunction<R> fun) {
       var levelsUp = 0;
       var shouldSkip = false;
+      var skippedObjectScope = false;
       for (var scope = this; scope != null; scope = scope.getParent()) {
         // for headers resolve variables one scope up
         if (scope instanceof EagerGeneratorScope) {
@@ -524,11 +525,24 @@ public final class SymbolTable {
             if (scope instanceof ObjectScope objectScope && objectScope.hasParams()) {
               levelsUp++;
             }
+            // An EagerGeneratorScope (for `when` predicates) skipped an ObjectScope.
+            // The ObjectScope may become a parse-time-invisible amend function at runtime,
+            // so for-gen variables at levelsUp == 0 need ReadFrameSlotNode
+            // (which calls skipInvisibleScopes) rather than ReadExactFrameSlotNode.
+            if (scope instanceof ObjectScope) {
+              skippedObjectScope = true;
+            }
             shouldSkip = false;
             continue;
           }
           var result = fun.apply(lex, levelsUp);
-          if (result != null) return result;
+          if (result != null) {
+            if (result instanceof ForGeneratorVariableOrLetBinding p && skippedObjectScope) {
+              //noinspection unchecked
+              return (R) new ForGeneratorVariableOrLetBinding(p.slot(), p.levelsUp(), true);
+            }
+            return result;
+          }
           if (scope instanceof MethodScope
               || scope instanceof ForGeneratorScope
               || scope instanceof LetExpressionScope) {
@@ -798,7 +812,7 @@ public final class SymbolTable {
         return null;
       }
       if (name.equals(binding.name())) {
-        return new ForGeneratorVariableOrLetBinding(binding.slot(), levelsUp);
+        return new ForGeneratorVariableOrLetBinding(binding.slot(), levelsUp, false);
       }
       return null;
     }
@@ -910,10 +924,10 @@ public final class SymbolTable {
     @Override
     public @Nullable VariableResolution doResolveProperty(String name, int levelsUp) {
       if (keyBinding != null && keyBinding.name().equals(name)) {
-        return new ForGeneratorVariableOrLetBinding(keyBinding.slot(), levelsUp);
+        return new ForGeneratorVariableOrLetBinding(keyBinding.slot(), levelsUp, false);
       }
       if (valueBinding != null && valueBinding.name().equals(name)) {
-        return new ForGeneratorVariableOrLetBinding(valueBinding.slot(), levelsUp);
+        return new ForGeneratorVariableOrLetBinding(valueBinding.slot(), levelsUp, false);
       }
       return null;
     }
diff --git a/pkl-core/src/main/java/org/pkl/core/ast/builder/VariableResolution.java b/pkl-core/src/main/java/org/pkl/core/ast/builder/VariableResolution.java
index 81c827b62..7c433478a 100644
--- a/pkl-core/src/main/java/org/pkl/core/ast/builder/VariableResolution.java
+++ b/pkl-core/src/main/java/org/pkl/core/ast/builder/VariableResolution.java
@@ -34,7 +34,8 @@ public sealed interface VariableResolution {
   // method, lambda, object body param
   record Parameter(int slot, int levelsUp) implements VariableResolution {}
 
-  record ForGeneratorVariableOrLetBinding(int slot, int levelsUp) implements VariableResolution {}
+  record ForGeneratorVariableOrLetBinding(int slot, int levelsUp, boolean needsFrameSkip)
+      implements VariableResolution {}
 
   // Implicit base module lookup
   record ImplicitBaseProperty() implements VariableResolution {}
diff --git a/pkl-core/src/test/files/LanguageSnippetTests/input/generators/forGeneratorNestedReference3.pkl b/pkl-core/src/test/files/LanguageSnippetTests/input/generators/forGeneratorNestedReference3.pkl
new file mode 100644
index 000000000..34dc2d5dd
--- /dev/null
+++ b/pkl-core/src/test/files/LanguageSnippetTests/input/generators/forGeneratorNestedReference3.pkl
@@ -0,0 +1,15 @@
+class Foo {
+  prop: String?
+}
+
+local foos: Mapping<String, Mixin<Foo>> = new Mapping<String, Mixin<Foo>> {
+  for (key, value in Map("foo", Map("key", "value"))) {
+    [key] = new {
+      when (value.keys.contains("key")) {
+        prop = value["key"]
+      }
+    }
+  }
+}
+
+res = foos["foo"].apply(new Foo {})
diff --git a/pkl-core/src/test/files/LanguageSnippetTests/output/generators/forGeneratorNestedReference3.pcf b/pkl-core/src/test/files/LanguageSnippetTests/output/generators/forGeneratorNestedReference3.pcf
new file mode 100644
index 000000000..d5bdd1d1d
--- /dev/null
+++ b/pkl-core/src/test/files/LanguageSnippetTests/output/generators/forGeneratorNestedReference3.pcf
@@ -0,0 +1,3 @@
+res {
+  prop = "value"
+}